I'm going to make a rather bold move and lump Royal penguins and Macaroni
penguins together. I think there's enough similarity between the two to make this
move since several now consider the Royal penguin to be a geographically
distinct race of Macaroni penguins. I'll talk about them together in this page.
    Macaroni penguins are easily identified from the other crested penguins:
their crests are bright orange and form on the centerline of the dorsal head and
point outwardly, slightly posteriorly. Royal penguins have the same configuration,
but they generally have white faces; Macaroni's have black faces. Males and females
of both penguins are monomorphic.
    Both Royals and Macaroni penguins weigh about 4.5 kilograms, but can
weigh more or less depending on the season. They reach about 70 cm in length. Numbers
vary slightly depending on whether you are talking about Royals; they reach about
65 cm in length.
    Together, Macaroni's and Royals comprise a population of about 10+ million
breeding pairs. Breaking it down, there are about 10 million pairs of Macaronis breeding
and about 115,000 pairs of Royals breeding. Thus, as you can see, Macaronis are more abundant
than Royals. However, their large number doesn't necessarily make them stable; Macaroni
penguins may be threatened by the increasing fishing industry. Royals are particularly
vulnerable to oil spills.
    Macaroni penguins inhabit the maritime
Antarctic regions and the Royal penguin inhabits the
Subantarctic regions. Some of the breeding ranges overlap: Macquarie Island. Macaronis
breed throughout South Shetland, South Sandwich, Bouvet Oy and Prince Edward Island. Niehter is
migratory and they both feed upon the same animals: crustaceans (euphausiids, amphipods),
fish and squid. They are both preyted upon by leopard seals, skuas, Wekas and petrels.
   
Macaroni penguins Eudyptes chrysolophus, which are probably
the largest members of the genus, approach the smaller pygoscelids in length
and weight. The orange-yellow crests originate in a broad band across the
forehead, and the chin and face are dark. Macaronis breed on South Georgia
and other islands of the Scotia Arc, on Bouvet Oy and Heard Island, and on
Marion Island, Iles Crozet and Kerguelen, and the Falkland Islands. No
subspecies have been postulated, although the Royal penguin on Macquarie
Island is by some authorities regarded as a subspecies of the Macaroni.
Macaronis form large, tightly packed colonies on flat or rough sloping
ground, often seeking the shelter of boulders or tussock grass. Notes on
their breeding appear in Matthews (1929) and Dowes et al. (1959),
though a comprehensive study of the species has yet to be made.
   
Royal penguins Eudyptes schlegeli are closely akin to Macaronis,
differing mainly in their possession of a predominantly grey or white face;
Carrick and Ingham (1970) and Warham ... give them only subspecific status.
They breed only on Macquarie Island, in large, dense colonies on
tussock-covered slopes of scree and beach. So densely packed are the
colonies that vegetation disappears and the birds incubate and rear their
chicks in mud. Preliminary accounts of long-term banding studies have
been published (Carrick and Ingham, 1967 and 1970), but little
information on the breeding and general biology of this species has yet
appeared.